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3.1 Mechanisms to inhibit proliferation and invasion of inflammatory synovial cells In RA pathology, FLS are key pathogenic cells that show proliferative and invasive phenotypes, secrete proteases and pro-inflammatory cytokines to damage cartilage and bone, are divided into multiple functional subtypes based on surface markers, possess phenotypic plasticity, and maintain inflammatory memory by interacting with T cells, B cells, and macrophages ( 2+ -dependent HDACs and NAD + -dependent HDACs