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Simultaneously, mechanical cues activates the Hippo pathway effector Yes-associated protein 1 (YAP), which translocates to the nucleus, where it promotes the transcription of profibrotic genes, reinforcing ECM deposition and stiffness (129)

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ROS act as secondary messengers to activate inflammasomes such as NLRP3, promoting the release of IL-1 and IL-18, and fostering the senescence-associated secretory phenotype (SASP)