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The L-cells in the intestinal epithelium express various nutrient-sensing receptors: Dietary fats are particularly potent GLP-1 secretagogues, activating free fatty acid receptors (such as GPR40 and GPR120) on L-cells Proteins and amino acids stimulate GLP-1 through peptide-sensing mechanisms and calcium-sensing receptors Carbohydrates trigger release via glucose transporters and sweet taste receptors expressed on enteroendocrine cells Dietary fibre , particularly soluble fibre, promotes GLP-1 secretion through fermentation products (short-chain fatty acids) produced by gut microbiota The rate and site of nutrient delivery also influences GLP-1 response

The "32mg" specification refers to the total peptide content held in the pre-filled pen reservoir, not the volume delivered per administration increment

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" Paracrine crosstalk between intestinal L- and D-cells controls secretion of glucagon-like peptide-1 in mice " Published in American Journal of Physiology - Endocrinology and Metabolism in 2019 Experimental proof for the paracrine regulation of intestinal GLP-1 secretion by somatostatin, providing background for the development of somatostatin-5-receptor antagonists for diabetes therapy