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The primary structure of a peptide usually is linear, but it also may be cyclic (following translation of the amino acid sequence, the N terminus and C terminus have connected via an amide bond) or branched (due to presence of residues with amino groups or carboxyl groups in their R chains) [6]

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The cyclization is formed via a lactam bridge between the side chains of Asp and Lys, constraining the peptide backbone into a stable, bioactive conformation